CHARACTER_STATEMENTS,STATE_1,STATE_2,STATE_3,STATE_(the columns to the right are also states),,,,
Ocellar corona,absent,present,,,,,,
Supraantennal groove or depression,absent,"present, accommodating scapes",,,,,,
Notch on medial margin of eye,absent,present,,,,,,
Position of toruli relative to eyes,below or level with lower margin of eyes,above lower margin of eyes,,,,,,
Position of toruli relative to clypeus,equal or closer to clypeus than its own diameter,further from clypeus than its own diameter,,,,,,
Subantennal shelf,absent,"present, antennal foramina facing upwards",,,,,,
Inner margin of torulus,not distinctly raised compared to outer margin,"with distinct projection, raised compared to outer margin",area between projections raised as well forming interantennal process,,,,,
Subantennal groove,absent or weakly developed,"present, distinct",,,,,,
Clypeus,not_inflected,inflected,,,,,,
Mandibular foramen,"absent, oral and mandibular foramen continuous",mandibular and oral foramina separated by subgenal sclerotisation,,,,,,
Occipital sulcus and ridge,absent,present,,,,,,
Position  occipital foramen,approx._halfway_between_top_of_head_and_oral_foramen,distance_from_top_of_head_to_occipetal_foramen_half_or_less_than_distance_to_oral_foramen,,,,,,
Position  posterior tentorial pits,adjacent/at level with occipital foramen/condyles,considerably_ventral_to_occipital_foramen,,,,,,
Sclerotization between occipital and oral foramina,"absent, foramina confluent","present, foramina separate",,,,,,
Ventral sclerotization configuration,"at most oblique ventral to tentorial pits, not extending anteriorly",horizontal ventral to pits and extending anteriorly,,,,,,
Postoccipital bridge,absent,bridge connecting insertion points of the ventral profurco-postoccipital muscles ventrally of the tentorial bridge present,,,,,,
Longitudinal sulci on ventral head sclerotization,none,"one median, at least ventrally","two sublateral, not merging ventrally",,,,,
Occipital carina,absent,present,,,,,,
Occipital carina configuration,reaching hypostome ventrally,not reaching hypostome and not continuous medially,continuous ventrally of occipital foramen,,,,,
Dorsal tentorial arm,"absent or reduced, not reaching head capsule","present, in contact with head capsule anteriorly",,,,,,
Shape  tentorial bridge,"broad, not above level of posterior tentorial arms","slender, arched above posterior tentorial arms",,,,,,
Corpotendon on tentorial bridge,absent,present anteriorly on tentorial bridge,,,,,,
Position of labrum,anterior,posterior,,,,,,
Shape of labrum,transverse,trapezoidal,thin and cylindrical,"long, thin and flattened","transverse, planar and weakly sclerotized to membranous",triangular,membranous sac,transverse with narrow medial projection
Labrum,present,absent,,,,,,
Exposure of labrum,exposed,concealed,,,,,,
Distal epipharyngeal wall,not synscleritors with labrum,synscleritors with labrum,,,,,,
Lateral epipharyngeal brush,absent,present,,,,,,
Epipharyngeal lobe,absent,present,,,,,,
Epipharyngeal setae,absent,present,dense medial patch of stout setae,,,,,
Tormae,"distinct apodemes continuous with labrum, not continuous medially","present, continuous medially",absent,,,,,
Sclerotization of labrum,heavily sclerotized,weakly scleritized,membranous,,,,,
Thickness of labrum,inner and outer surfaces separate,flaplike,,,,,,
Labral setation,scattered setae over anterior surface,setae restricted to apical margin,bare,,,,,
Labral apodemes,absent,"present, long and thick","present, short, thin and articulating with head capsule",,,,,
Sitophore,absent,"present, does not extend distally of the functional mouth","present, extends distally of the functional mouth",,,,,
Sitophore sensillae,absent,patch (>4) of lateral sensilla present,1-2 lateral sensilla only,,,,,
Hypopharyngeal lobe,absent,present,,,,,,
Infrabuccal pouch,small and/or with inconspicuous armature,"large, with distinct cuticular teeth developed for grinding pollen",forming a distinct gnathal pouch,,,,,
Hypopharyngeal pectens,absent,minute hairs on hypopharyngeal rods,distinct brushes,,,,,
Left mandible dentition,no teeth,one extra tooth,two extra teeth,four or more distinct teeth,,,,
Dentition of right mandible relative to left,one_less,equal_number,,,,,,
Mola on right mandible,present,absent,,,,,,
Mandibular peg on ventral margin,absent,present,,,,,,
Mandibular rods glands,absent,present,,,,,,
Lateral articulation of mandible,with_distinct_posterolateral_condyle,truncate_connection,medial_condyle,anterolateral_condyle,,,,
Lateral mandibular muscle attachment,hidden,visible_externally,,,,,,
Shape of glossa  alaglossa ,lobate,flat,folded into an inverted U-shape,hoodlike_and_folded_medially,semispherical,absent_or_very_reduced,,
Setation of glossa,scattered_setae,distinct_rows_of_flattened_setae,bare,densely_setose,spinose,,,
Glossal fringe,absent,present,,,,,,
Submarginal setae of glassa,absent,present,,,,,,
Campaniform sensillae,absent,present,,,,,,
Acroglossal button,absent,present,,,,,,
Glossal margin,entire,slightly_emarginate,deeply_incised,,,,,
Paraglossae,"club-shaped, without microtrichia",lobate with setae or flattened scale-like microtrichia in rows,flattened with scale-like microtrichia in transverse rows,distal scale-bearing part reduced,"totally reduced, fused with glossa",,,
Paraglossal basal setation,setose_or_bare,spinose,,,,,,
Number of labial palp segments,0,2,4,,,,,
Rod like sensilla,absent,specialised_sensilla_on_distal_labial_palp_segment_present,"present, situated in invagination with pore opening",,,,,
Subapical sensilla,absent,present,,,,,,
Postmentum,absent,faint_trace,,,,,,
Number of maxillary palp segments,0,2,4,6,,,,
Maxillary palp shape,"thin, not leg-like","thick, leg-like, with a setae-bearing distal part",,,,,,
Sensillum on third maxillary palpomere,absent,present,,,,,,
Third maxillary palpomere,cylindrical,"enlarged, broader than the following palpomere, and triangular",,,,,,
Shape of lacinia,rounded,acuminate,,,,,,
Galea shape,entire,divided_and_broad,divided_and_longitudinally_elongate,absent,,,,
Galea division,with_two_apices,unipartite,,,,,,
Stipes sensillum,absent,single_prominent_sensillum,,,,,,
Pronotum shape anteriorly,protruding,reduced,,,,,,
Pronotum  anteroventral longitudinal carinae,absent,two_parallel_carinae_with_concave_area_inbetween_present_,,,,,,
Pronotum  submedian depressions,absent,pits_corresponding_to_internal_concavities_present,,,,,,
Pronotum  length,long_dorsally_of_transverse_sulcus,short_dorsally_of_transverse_sulcus,,,,,,
Pronotum orientation,pronotum_posteriorly_of_transverse_sulcus_oblique,pronotum_posteriorly_of_transverse_sulcus_vertical,,,,,,
Pronotum  transverse carina,absent,"present, traversing pronotum dorsally of transverse sulcus",,,,,,
Pronotum  anterior surface,"convex or straight, not accommodating head","concave, smooth, delimited by lateral projections, accommodating head",,,,,,
Pronotum  dorsomedially,without_shelf,with_smooth_shelf_accommodating_prophragma,,,,,,
Pronotum  submedian apodemes,absent,small_apodemes_present,,,,,,
Pronotum  anterolateral corners,rounded,with_distinct_notch_accommodating_anterolateral_corner_of_mesoscutum,,,,,,
Pronotum  anterolateral internal carina,absent_or_weakly_developed,distinct_transverse_carina_present_anterolaterally,,,,,,
Pronotum  posterodorsal corner,without_notch,"with notch above anterior spiracle, accommodating tegula",,,,,,
Pronotum  posterolateral margin,with_incurvation_accommodating_anterior_spiracle,straight,with_extended_lobe,,,,,
Pronotum  anterior spiracle,"visible, posterior to posterolateral margin of pronotum",covered_by_posterolateral_margin_of_pronotum,"visible, surrounded by continuous cuticle",,,,,
Pronotum  mesopleural attachment,"movable relative to mesopleuron, or not abutting",rigidly_attached_to_mesopleuron,,,,,,
Pronotum  lateroventral corners,not_projecting_medially,with_medial_inflexions_abutting_mesopleuron,"fused, forming bridge behind procoxae",,,,,
Prepectus,absent,independent_sclerite_present,,,,,,
Prepectus,exposed,concealed,,,,,,
Prepectus  pronotal inflection,absent,present,,,,,,
Prepectus inflection  dorsal extent,not_extending_dorsally_of_anterior_spiracle,extending_dorsally_of_anterior_spiracle,,,,,,
Prepectus inflection  ventral configuration,separate_or_at_most_slightly_abutting,broadly_abutting,fused_together_or_with_prospinasternum,,,,,
Position of occlusor muscle apodeme,midway on posterolateral pronotal margin,ventrally on posterolateral pronotal margin,,,,,,
Position  anterior thoracic spiracle,more_than_one_spiracular_diameter_from_posterodorsal_corner_of_pronotum,at_posterodorsal_corner_of_pronotum,dorsally_between_pronotum_and_mesoscutum,,,,,
Projection ventrad anterior spiracle,absent,narrow projection ventrally of spiracle present,,,,,,
Propleura  cervical prominences,"not retracted, cervical swellings exposed","retracted, swellings concealed",,,,,,
Propleura  position of cervical prominences,at_anterodorsal_corners_of_propleura,ventrally_of_anterodorsal_corners_of_propleura,,,,,,
Propleuron  cervical apodeme,entirely_absent,"short, inconspicuous","well developed, distinct",,,,,
Propleuron  cervical apodeme configuration,separate_from_propleura_posteriorly,curves_laterally_to_fuse_with_propleura_posteriorly,,,,,,
Propleuron  cervical lines,laterocervicalia_and_propleuron_proper_at_most_articulating,"laterocervicalia and propleuron partly fused, cervical line present","laterocervicalia and propleuron completely fused, cervical line absent",,,,,
Propleura  dorsal part,"not inflected, sculpture similar to rest of propleura",inflected_with_smooth_or_shagreened_sculpture,,,,,,
Dorsal margins of propleura,widely separated for most of their length,abutting for at least half their length,,,,,,
Propleuron  propleural arm,absent,present,,,,,,
Configuration of propleural arms,"erect, not abutting profurca posteriorly","horisontal, extending medially posteriorly of profurca",,,,,,
Propleura  lateroventral carina,absent,longitudinal_carina_on_lateroventral_margin_of_propectus_present,,,,,,
Propleura  median margins,widely_separate_for_their_entire_length,"diverging posteriorly, prosternum broadly exposed","abutting for most of their length, prosternum partly concealed",,,,,
Propleura  epicoxal lobes,posterolateral_corner_of_propleuron_not_extended,"extended as epicoxal lobe, covering procoxa proximally",,,,,,
Propleura  posterolateral groove,absent,"distinct groove dorsally on corners, articulating with pronotum",,,,,,
Propleura  prothoracic katepisterna,absent,present,,,,,,
Propleuron  katepisternum configuration,abutting_propleuron_at_articulation_with_procoxa,separate_from_propleuron,,,,,,
Prosternum  anteroventral part,at_most_extending_slightly_anterior_to_posterior_margin_of_propleura,extending_well_anterior_to_posterior_margin_of_propleura,,,,,,
Prosternal spine,absent,median posteriorly directed spine present,,,,,,
Prosternum  laterosternal sclerites,absent,present,,,,,,
Prosternum  subdivision,not_subdivided,transversely_subdivided_dorsally_by_deep_lateral_incisions,,,,,,
Profurca  number of profurcal pits,one,two,,,,,,
Profurca  anterior apodemes,absent_or_weakly_developed,"well developed, continuous with tendons",,,,,,
Profurca  bridge,absent,"present, low","present, high and curving posteriorly",,,,,
Profurca  articulation with propleuron,lateral_profurcal_arm_not_articulating_with_propleuron,lateral_profurcal_arm_articulating_with_propleuron_posterodorsally,,,,,,
Foreleg  procoxa,"broad basally, proximal foramen width at least half maximal width of procoxa","constricted basally, proximal foramen width less than half maximal width",,,,,,
Foreleg  procoxal transverse carina,absent,developed_for_at_least_part_of_coxa,,,,,,
Foreleg  protibial furrow,absent,U-shaped furrow present,,,,,,
Foreleg  calcar,anterior_apical_protibial_spur_unmodified,anterior_apical_protibial_spur_modified_into_calcar_for_antenna_cleaning,,,,,,
Foreleg  posterior apical protibial spur,absent_or_reduced,present,,,,,,
Foreleg  probasitarsal notches,absent,present,,,,,,
Foreleg  probasitarsal combs,absent_or_weakly_developed,"present, well developed",,,,,,
Foreleg  probasitarsal spur,absent,present,,,,,,
Mesonotum  prophragma,absent_or_reduced,well_developed,,,,,,
Mesonotum  prophragma shape,bilobed,not_subdivided_by_slit,,,,,,
Mesonotum  prophragma laterally,"not extended, higher submedially than laterally","extended, higher laterally than submedially",,,,,,
Mesonotum  anterior mesoscutal sulcus,absent,transverse_groove_accommodating_dorsal_margin_of_pronotum_present,,,,,,
Mesonotum  anterior mesoscutal sulcus configuration,at_most_extending_slightly_onto_prophragma_anteromedially,with_distinct_ventral_extension_onto_prophragma_anteromedially,,,,,,
Mesonotum  median mesoscutal sulcus and ridge,well_developed_internally_and_externally,developed_only_externally,absent_or_not_fully_developed_posteriorly,,,,,
Mesonotum  anteroadmedian signum,absent,present,,,,,,
Mesonotum  notauli,well_developed_internally_and_externally,developed_only_externally,"absent, at least posteriorly",,,,,
Mesonotum  notauli configuration,"separate, do not join prior to reaching transscutal fissure",converge_medially_to_join_prior_to_reaching_fissure,"merge into U-shaped sulcus, not reaching fissure",,,,,
Mesonotum  parapside,absent,present,,,,,,
Mesonotum  parascutal carina,absent,area_accommodating_tegula_delimited_by_longitudinal_carina_present,,,,,,
Mesonotum  transscutal articulation,absent,"present, not less sclerotised than adjacent cuticle","present, less sclerotised than adjacent cuticle",,,,,
Mesonotum  transscutal carina,absent,present,,,,,,
Mesonotum  size of dorsal axillar surface,"large, mesoscutellar sulcus U- or V-shaped","small, mesoscutellar sulcus straight medially",,,,,,
Mesonotum  axillar carina,absent,longitudinal_carina_extending_dorsally_from_scutellar_arm_present,,,,,,
Mesonotum  lateral corners of axilla,"rounded, no distinct projection dorsally",axillar_carina_with_distinct_posterior_projection_dorsally,,,,,,
Mesonotum  axillar phragma,absent,present,,,,,,
Mesonotum  scutoscutellar sulcus,different_configuration_than_state_1,consisting_of_one_or_two_wide_depressions_with_small_pits_laterally,,,,,,
Mesonotum  mesoscutellar carina,absent_or_weakly_developed_medially,"present, distinct throughout",,,,,,
Mesonotum  configuration of mesoscutellar carina,reaching transscutal carina/articulation medially,not reaching transscutal carina/articulation,,,,,,
Mesonotum  internal median longitudinal mesoscutellar carina,absent,present,,,,,,
Mesonotum  internal septum,absent,septum_with_small_fenestrum_spanning_at_least_part_of_mesoscutellum_ventrally,,,,,,
Mesonotum  longitudinal subdivision of scutellum,not_subdivided,subdivided_by_submedian_longitudinal_grooves,,,,,,
Mesonotum  frenum,mesoscutellum_not_subdivided,"mesoscutellum transversely subdivided by line, fold, or carina",,,,,,
Mesonotum  mesoscutellum posteriorly,not_as_state_1,medially_with_row_of_sculptured_depressions,,,,,,
Mesonotum  mesoscutellar appendage,absent_or_weakly_developed,projection_separated_from_scutellum_by_distinct_line_present,,,,,,
Mesonotum  mesoscutellar arm configuration,"narrow, erect, slender","broad, low, smooth",,,,,,
Mesonotum  origin of mesoscutellar arms,at_posterolateral_corners_of_mesoscutellum,anterolaterally_on_mesoscutellum,,,,,,
Mesonotum  mesoscutellar arm distally,not_extending,distinctly_extended_laterally,,,,,,
Mesophragma  pseudophragmal lobe s ,absent_or_weakly_developed,"well developed, distinct lobes",,,,,,
Mesophragma  anterior margin position,at_most_displaced_slightly_posterior_to_posterior_margin_of_mesoscutellum,displaced_posteriorly_into_propodeum_for_at_least_the_length_of_the_mesoscutellum,,,,,,
Mesophragma  mesophragmal pockets,"weakly developed, no curved line anterodorsally on mesophragma","well developed, with deeply curved posterior margin",,,,,,
Mesophragma  median longitudinal carina,absent_or_weakly_developed,distinct_carina_extending_ventrally_for_most_of_phragma,,,,,,
Mesophragma  mesolaterophragmal lobes,reduced_or_absent,"present, well developed",,,,,,
Mesophragma  number of mesolaterophragmal lobes,one,two,,,,,,
Mesophragma  mesolaterophragmal apodeme shape,"short, rounded","elongate, slender",,,,,,
Mesophragma  mesopostnotum,exposed,concealed_by_anterior_part_of_metanotum,,,,,,
Mesophragma  lateral attachment of mesopostnotum,"broad, exposed","invaginated, but still developed","reduced, mesopostnotum entirely internal",,,,,
Mesophragma  lateral flanges,absent,distinct_flanges_present_anteromedially,,,,,,
Mesopectus  configuration with mesonotum,"not broadly abutting, tegulum and pronotum adjacent","abutting for considerable distance, tegulum well separated from pronotum",,,,,,
Mesopectus  tegula configuration,exposed,concealed_by_dorsolateral_corner_of_pronotum,,,,,,
Mesopleuron  external part of basalar,not_enlarged,"conspicuously enlarged, extending ventrally of anterior spiracle",,,,,,
Mesopleuron  basalar apodeme,"large, cup-shaped","reduced, tendon-like",,,,,,
Mesopleuron  postspiracular sclerite,absent,present,,,,,,
Mesopleuron  posterior thoracic spiracle configuration,"exposed, in concavity in mesopleural margin","exposed, above mesopleural margin",not_visible_externally,,,,,
Mesopleuron  supramesopleural sclerite,absent,sclerite_between_mesopleuron_and_metapleural_arm_present,,,,,,
Mesopleuron  mesepimeral ridge,absent_or_weakly_developed,distinct_ridge_extending_from_dorsal_margin_of_mesopleuron_to_approx._halfway_to_mesocoxal_articulation,distinct_ridge_extending_from_dorsal_margin_of_mesopleuron_to_mesocoxal_articulation,,,,,
Mesopectus  mesopleural ridge,absent,ridge_extending_between_anterior_margin_of_mesopleuron_and_mesocoxal_articulation,,,,,,
Mesopleuron  mesopleural pit,absent,small_pit_present,,,,,,
Mesopleuron  mesopleural apodeme,absent_or_weakly_developed,apodeme_present,,,,,,
Mesopleuron  apodeme configuration,separate_from_mesepimeral_ridge,continuous_with_mesepimeral_ridge,,,,,,
Mesopectus  mesopleural triangle,at_most_slight_depression_present_dorsally_on_mesopleuron,distinct_triangular_depression_extending_across_dorsal_mesopleuron,,,,,,
Mesoplueron  prospinasternum,separate_sclerite,fused with mesopleuron and/or pronotum,,,,,,
Mesopleuron  prospinasternal apodeme,absent,"present, well developed",,,,,,
Mesopleuron  acetabular epicnemial carina,absent_or_weakly_defined,"distinct, extends below procoxae",,,,,,
Mesopleuron  procoxal concavities,absent_or_weakly_developed,concavities_accommodating_procoxae_present,,,,,,
Mesopleuron  subpleural signum,absent,longitudinal_line_or_crease_present,,,,,,
Mesopectus  pseudosternal sulci,absent_or_not_developed_internally,well_developed_internally_and_externally,,,,,,
Mesopectus  sternaular carina,absent_or_weakly_developed,oblique_internal_carina_on_mesepisternum_present,,,,,,
Mesopleuron  posterior transverse carina,absent,"present, delimiting concave or smooth area just anteriorly of mesocoxal foramina",,,,,,
Mesopleuron  mesocoxal foramina,"widely open dorsally, not surrounded by mesopleural cuticle","foramen constricted, but still with narrow opening dorsally","closed dorsally, surrounded by continuous mesopleural cuticle",,,,,
Mesopleuron  median mesocoxal articulation,absent,"present, on projection",,,,,,
Mesopleuron  medial mesocoxal articulation position,"situated proximally, articulations not elongated","situated distally, on medial surface of mesocoxa",,,,,,
mesofurca  mesofurcal pit,situated_at_level_with_mesocoxal_foramina,situated_anteriorly_of_mesocoxal_foramina,,,,,,
Mesofurca  mesospina and metafurco mesospinal muscle,absent,"present, short, erect","elongate, projecting posteriorly between metafurcal arms",,,,,
Mesofurca  arms proximally,with_at_most_shallow_depressions,with_distinct_concavities_dorsally,,,,,,
Mesofurca  anterior arm,"short, inconspicuous",elongate,,,,,,
Mesofurca  bridge  R99  54 ,absent,present,,,,,,
Mesofurca  anterior projection on bridge,absent,present,,,,,,
Mesofurca  lateral arm,"short, not extending towards mesopleuron","elongate, extending towards mesopleuron",,,,,,
Midleg  mesocoxa,not_subdivided,subdivided,,,,,,
Midleg  proximal part of mesocoxa,well_developed,very_reduced,,,,,,
Midleg  coxo trochanter articulation,"broad, trochanter cylindrical proximally","narrow, trochanter with flattened disk proximally",,,,,,
Midleg  trochantellus,absent,"femur subdivided proximally, forming small trochantellus",,,,,,
Metanotum,"reduced, antecostal sulcus adjacent to mesoscutellum","well developed, antecostal sulcus well separated from mesoscutellum",,,,,,
Metanotum  orientation,"more or less horisontal, angled less than 90o relative to mesoscutellum","vertical, angled 90o relative to mesoscutellum",,,,,,
Metanotum  cenchri,absent,present,,,,,,
Metanotum hindwing tegula,absent,present,,,,,,
Metanotum  humeral sclerite,"absent, anterior metanotal wing process continuous with remainder of metanotum",anterior_metanotal_wing_process_situated_on_separate_sclerite,,,,,,
Metanotum  metascutellum,absent_or_reduced,"present, well developed",,,,,,
Metanotum  metascutellum configuration,"reaching anterior margin of metanotum, abutting mesoscutellum",not_reaching_anterior_margin_of_metanotum,,,,,,
Metanotum  incurvation posterad metascutellum,at_most_weakly_developed,"present, distinct",,,,,,
Metanotum  lateral longitudinal carina,"absent or reduced, not traversing metanotum","well developed, traversing entire metanotum",,,,,,
Metanotum  lateral incision,absent,anterolateral_membraneous_area_present,,,,,,
Metapleuron  apodemal part of metabasalare,absent_or_reduced,"present, independent cup-shaped sclerite","present, continuous with metapleuron",,,,,
Metapleuron  metapleural arm   T1 association,not_abutting_T1,abutting_T1,fused_with_T1,,,,,
Metaplueron  metaplural arm configuration,continuous_with_rest_of_metapleuron,separated_from_rest_of_metapleuron_by_anterolateral_part_of_T1,,,,,,
Metapleuron  metapisternal anapleural cleft,absent,present,,,,,,
Metapleuron  metepimeron   T1 association,at_most_fused_for_short_distance,broadly_fused,,,,,,
Metapluron  metepimeron,well_developed_throughout,reduced_posteriorly,,,,,,
Metapleuron  articular inflection,absent,present,,,,,,
Metapleuron  dorsal longitudinal ridge,absent,distinct_ridge_ventral_to_propodeal_spiracle_present,,,,,,
Metapleuron  metapleural pit,entirely_absent,at_least_shallow_depression_present,,,,,,
Metapleuron  metapleural apodeme,absent_or_weakly_developed,"present, well developed",,,,,,
Metapleuron  mesopleural metapleural association,"might articulate or abut ventrally, but not fused",fused_ventrally,fused_ventrally_and_laterally,,,,,
Metapleuron  paracoxal notches,absent,present,,,,,,
Metapleuron  median longitudinal carina projection,absent,"present, separating metepisternal depressions",depressions_separated_by_broad_swelling,,,,,
Metapleuron  lateral longitudinal carina,absent,low_carina_laterally_of__metepisternal_depressions_present,projections_at_anterolateral_corners_of_metepisternal_depressions_present,,,,,
Metapleuron  ventral transverse carina,absent_or_weakly_developed,transverse_carina_anterior_to_metacoxal_foramina_present,,,,,,
Metapleuron  area anterior to paracoxal ridge sulcus,reduced_medially,well_developed,,,,,,
Metapleuron  anterior paracoxal sulci ridges,absent,present,,,,,,
Metapleuron  posterior paracoxal ridge,absent,present,,,,,,
Metapleuron  anterior paracoxal ridge,not_reaching_metapleural_apodemes,extending_to_metapleural_apodemes,"continuous with marginal metapleural apodeme, separate from metapleural apodeme",,,,,
Metapleuron  metapectal plates,absent,low_lobe_present_laterally_on_anterior_paracoxal_ridge,,,,,,
Metapluron  metacoxal foramina,well_separated_from_mesocoxal_foramina,"adjacent to mesocoxal foramina, ventral metepisternum reduced",,,,,,
Metapleuron  median coxal articulation,"reduced, at most developed as low  swelling",articulation_point_well_developed,,,,,,
Metapleuron  median coxal articulation position,"ventromedial to lateral articulation, axis of movement oblique","approx. ventral to lateral articulation, axis of movement vertical",,,,,,
Metapleuron  lateral coxal articulation,"exposed, visible laterally",concealed_in_lateral_view,,,,,,
Metapleuron  articulation notch,absent_or_weakly_developed,notch_in_foraminal_rim_at_metacoxal_lateral_articulation_well_developed,,,,,,
Metafurca  metafurcal pit position,closer_to_metacoxal_foramina_than_to_anterior_margin_of_metapectus,closer_to_anterior_margin_of_metapectus_than_to_metacoxal_foramina,,,,,,
Metafurca  number of metafurcal pits,one,two,,,,,,
Metafurca  position,metafurcal_arms_arise_posteriorly_on_metadiscrimen,metafurcal_arms_arise_at_or_close_to_anterior_end_of_metadiscrimen,,,,,,
Metafurca  furca discrimen configuration,"adjacent, arise from discrimen","widely separate, not converging, detached from discrimen",,,,,,
Metafurca  connection to mesofurca,separate_from_mesofurca,fused_with_mesofurca_for_some_distance,,,,,,
Metafurca  anterior metafurcal arm,"elongate, at most slightly shorter than lateral arm","reduced, considerably shorter than lateral arm",,,,,,
Metafurca  lateral metafurcal arm,short,"elongate, extending towards metapleural apodeme",fused_with_metapleural_apodeme,,,,,
Hind leg  trochantins,absent,present,,,,,,
Hind leg  size of coxa,"short, less than 3/4 length of hindfemur+trochanter","elongate, more than 3/4 length of hindfemur+trochanter",,,,,,
Hind leg  femoral spines,absent_or_weakly_developed,at_least_one_distinct_spine_present_ventrally,,,,,,
Hind leg  tibial preapical spurs,absent,present,,,,,,
Hind leg  tibial brush,absent,dense_brush_of_setae_present_on_at_least_part_of_median_side,,,,,,
Hind leg  tarsal plantulae,absent,present,,,,,,
Propodeum T1 medially,divided,continuous,,,,,,
Propodeum  metapostnotum association,"separated by unsclerotised line, connection movable",immovably_fused,,,,,,
Propodeum  median antecostal projections,absent,one_projection_present_anteriorly_and_posteriorly_of_antecosta_,several_projections_present_anteriorly_and_posteriorly_of_antecosta_,,,,,
Propodeum  lateral antecostal projections,absent,present_anteriorly_and_posteriorly_of_antecosta,,,,,,
Propodeum  metaphragma,absent,present_only_laterally,"present, continuous medially",,,,,
Propodeum  metalaterophragmal lobes,weakly_developed_or_absent,well_developed,,,,,,
Propodeum  metaphragma metapleural apodeme configuration,separate,metaphragma_and_metapleural_apodeme_fused,,,,,,
Propodeum  metapostnotum,continuous_medially,subdivided_medially,,,,,,
Propodeum  median projection,absent,distinct_projection_present_medially,,,,,,
Propodeum  median longitudinal carina,absent,"present, well developed",,,,,,
Propodeum  lateral longitudinal carina,absent,distinct_ridge_dorsal_to_propodeal_spiracle_present,,,,,,
Propodeum  propodeal spiracle configuration,"elongate, slit-like",rounded,,,,,,
Propodeum  propodeal spiracle laterally,exposed,covered_by_flap,,,,,,
Propodeum  spiracle position  horizontal ,not_further_than_halfway_between_antecosta_and_T1_posterior_margin,closer_to_T1_posterior_margin_than_to_antecosta,,,,,,
Propodeum  spiracle position  vertical ,not adjacent to propodeal-metapleuron boundary,abutting_boundary,,,,,,
Propodeum  transverse carina,absent,distinct_carina_present_some_distance_from_propodeal_foramen,,,,,,
Propodeum  propodeal foramen,"not developed, T2 broadly attached to T1",T1_articulating_with_T2_but_not_constricted_laterally,"T1 posteriorly constricted laterally, articulating with T2",,,,,
Propodeum  propodeal foramen configuration,continuous_with_metacoxal_foramina,separated_from_metacoxal_foramina_by_sclerotised_bridges,,,,,,
Propodeum  position of propodeal foramen,situated_posteriorly_of_anterior_margin_of_metacoxal_foramen,situated_at_level_with_metacoxal_foramen,"dorsally, at least halfway between metacoxal foramina and metapostnotum","adjacent to antecostal sulcus, compressing sulcus medially",,,,
Propodeum  propodeal foramen ventrally,straight,distinctly_incurved,,,,,,
Propodeum  propodeal foramen shelf,absent,distinct_shelf_extending_ventrally_into_body_cavity_from_foramen,,,,,,
Propodeum  articulating teeth,absent_or_weakly_developed,distinct_teeth_present,,,,,,
First abdominal sternum,absent or reduced,present,,,,,,
Petiole  T2 and S2 configuration,separate,"fused, at least anteriorly",,,,,,
Petiole  posteriorly,wider_than_anterior_foramen,"narrow, approx. same width as anterior foramen",,,,,,
Petiole  articulating condyle,"reduced or absent, petiolar foramen subcircular","well developed, T-shaped in anterior view",,,,,,
Petiole  anterior width,"not broad, condyle occupying at least 1/2 of anterior width","broad, condyle less than 1/3 of anterior width",,,,,,
Petiole  transverse carina on T2,absent,"present, situated just posteriorly of articulation",,,,,,
Petiole  T2 longitudinal internal carina,absent,"carina present posteriorly, externally indicated by furrow",,,,,,
Petiole  S2,at_least_partly_sclerotised,membraneous_medioventrally,,,,,,
Petiole  S2 posteriorly,equally_sclerotised_or_membraneous_throughout,less_heavily_sclerotised_posteriorly_for_at_least_half_the_length_of_T2,,,,,,
Petiole  transverse carina anteriorly,absent,transverse_carina_posteriorly_of_sensillar_patches_present,,,,,,
Petiole  longitudinal carina anteriorly on S2,absent,present_posteriorly_of_anterior_margin,,,,,,
Muscles  median pronoto prophragmal muscle,absent,present,,,,,,
Muscles  site of origin of the dorsal pronoto procoxal muscle,from_along_the_dorsal_margin_of_the_pronotum,distinctly_ventrally_of_the_dorsal_margin_of_the_pronotum,from_the_ventral_surface_of_the_anterior_thoracic_spiracle_occlusor_muscle_apodeme,,,,,
Muscles  ventral pronoto procoxal muscle,absent,present,,,,,,
Muscles  pronoto mesobasalar muscle,absent,present,,,,,,
Muscles  pronoto profurcal muscle,absent,present,,,,,,
Muscles  pronoto third axillary sclerite of forewing muscle,absent,present,,,,,,
Prepectus  anterior spiracle occlusor muscle,arise_from_free_prepectus,arise_from_mesopleuron,arise_from_pronotum,,,,,
Muscles  site of pronotal origin of the anterior thoracic spiracle occlusor muscle,originates_at_least_partly_from_the_occlusor_muscle_apodeme,originates_from_the_wall_of_the_pronotum,,,,,,
Muscles  lateral metapleuro postoccipital muscle,absent,present,,,,,,
Muscles  propleural arm postoccipital muscle,absent,present,,,,,,
Muscles  propleural arm protrochanteral muscle,absent,present,,,,,,
Muscles  mesofurco propleural arm muscle,absent,present,,,,,,
Muscles  profurco prophragmal muscle,absent,present,,,,,,
Muscles  origin of median profurco procoxal muscle,from_the_posteroventral_part_of_the_profurca,from_the_ventral_part_of_the_propleural_arm,,,,,,
Muscles  profurco protrochanteral muscle,absent,present,,,,,,
Muscles  prospinasterno profurcal muscle,absent,present,,,,,,
Muscles  dorsal mesofurco profurcal muscle,absent,present,,,,,,
Muscles  prospinasterno procoxal muscle origin,from_independent_prospinasternum,from_anterior_mesopleural_margin,from_ventral_pronotal_sclerotisation,from_independent_prepectus,,,,
Muscles  mesonoto mesobasalar muscle,absent,present,,,,,,
Muscle  mesonoto mesolaterophragmal muscle,absent,present,,,,,,
Muscles  mesonoto mesocoxal muscles,absent,present,,,,,,
Mescles  anterior mesonoto mesotrochanteral muscle,absent,present,,,,,,
Muscles  posterior mesonoto mesotrochanteral muscle,absent,present,,,,,,
Muscles  anterior mesonoto metanotal muscle,absent,present,,,,,,
Muscles  posterior mesonoto metanotal muscle,the_two_bands_of_the_paired_muscle_inserts_medially_on_the_metanotum,the_two_bands_of_the_paired_muscle_inserts_laterally_on_the_metanotum,,,,,,
Muscles  site of origin of second mesopleuro mesonotal muscle,from_the_mesopleural_ridge_or_posterior_to_the_mesopleural_ridge_from_the_mesopleural_arm_,anteriorly_of_the_mesopleural_ridge_or_from_the_wall_of_the_mesopleuron_sometimes_at_least_partly_from_the_mesopleural_apodeme,,,,,,
Muscles  intersegmental membrane mesobasalar muscle,absent,present,,,,,,
Muscles  mesocoxomesobasalar muscle,absent,present,,,,,,
Muscles  mesotrochantero mesobasalar muscle,absent,present,,,,,,
Muscles  mesopleuro mesosubalar muscle,absent,present,,,,,,
Muscles  mesocoxo mesosubalar muscle,absent,present,,,,,,
Muscles  mesofurco mesopleural muscle,absent,present,,,,,,
Muscles  site of origin of the mesopleuro mesocoxal muscle,originates_from_the_anterior_surface_of_the_mesopleural_ridge,originates_from_the_wall_of_the_mesopleuron_sometimes_partly_from_the_mesopleural_apodeme,,,,,,
Muscles  mesopleuro third axillary sclerite of hind wing muscle,absent,present,,,,,,
Muscles  posterior mesofurco mesolaterophragmal muscle,absent,present,,,,,,
Muscles  lateral mesofurco mesotrochanteral muscle,absent,present,,,,,,
Muscles  medial mesofurco mesotrochanteral muscle,absent,present,,,,,,
Muscles  site of origin of the median mesofurco mesotrochanteral muscle,originates_submedially_from_the_lateral_part_of_the_mesofurcal_arm,originates_from_the_mesofurcal_bridge_or_from_the_anterior_mesofurcal_arm,,,,,,
Muscles  mesofurco metabasalar muscle,absent,present,,,,,,
Muscles  mesospinasterno metafurcal muscle,absent,present,,,,,,
Muscles  metanoto metabasalar muscle,absent,present,,,,,,
Muscles  median metapleuro metanotal muscle,absent,present,,,,,,
Muscles  site of origin of the median metapleuro metanotal muscle,originates_anteriorly_of_the_paracoxal_ridge,originates_posteriorly_of_the_paracoxal_ridge,,,,,,
Muscles  metalaterophragmo metafurcal muscle,absent,present,,,,,,
Muscles  metanoto metalaterophragmal muscle,absent,present,,,,,,
Muscles  anterior metanoto metacoxal muscle,absent,present,,,,,,
Muscles  posterior metanoto metacoxal muscle,absent,present,,,,,,
Muscles  metanoto metatrochanteral muscle,absent,present,,,,,,
Muscles  site of origin of the anterior metapleuro metabasalare muscle,"from anteriorly of the paracoxal ridge, but posterior to the anterior margin of the metapleuron",from the wall of the fused meso-metapectus,,,,,,
Muscles  lateral metapleuro metabasalar muscle,absent,present,,,,,,
Muscles posterior metapleuro metabasalare muscle,absent,present,,,,,,
Muscles  metacoxo metabasalar muscle,absent,present,,,,,,
Muscles  metacoxo metasubalar muscle,absent,present,,,,,,
Muscles  anterior metapleuro metafurcal muscle,absent,present,,,,,,
Muscles  posterior metapleuro metafurcal muscle,absent,present,,,,,,
Muscles  lateral metapleuro metacoxal muscle,"originates from the metapleural apodeme and sometimes from the metapleuron ventrally of the metapleural apodeme, site of origin",originates_exclusively_from_the_paracoxal_ridge,,,,,,
Muscles  site of origin of the lateral metapleuro metacoxal muscle,does_not_extend_to_the_propodeum,extends_to_the_propodeum,,,,,,
Muscles  metapleuro propodeal muscle,absent,present,,,,,,
Muscles  metasterno second abdominal sternal muscle,absent,present,,,,,,
Muscles  site of origin of the metapleuro metatrochanteral muscle,does_not_extend_to_the_propodeum,extending_to_the_propodeum,,,,,,
Muscles  second abdominal sternal metacoxal muscle,absent,present,,,,,,
Muscles  propodeo second abdominal segment muscle,absent,present,,,,,,
Muscles  site of origin of the dilator muscle of the propodeal spiracle,originates_just_dorsally_of_the_lateral_metacoxal_articulation,"originates from the metapleural apodeme, anteriorly of the propodeal spiracle",originates_from_the_propodeum_on_the_level_of_the_propodeal_foramen,,,,,
Forewing  sculpture of retinaculum,with_microspicules,with_crenulae,smooth,,,,,
Forewing  costal and radial vein fusion,not_fused,fused_along_their_lengths,,,,,,
Forewing  cubital  Cu  vein shape,"bent sharply towards the posterior margin, apical to 1m-cu crossvein","not bent sharply towards the posterior margin, apical to 1m-cu crossvein",,,,,,
Forewing  costal  C  vein presence,complete_as_a_tubular_vein,absent_or_present_but_not_complete_as_a_tubular_vein,,,,,,
Forewing  stigma presence,present,absent,,,,,,
Forewing  subcostal  SC  vein presence,present_as_a_longitudinal_vein,present_as_an_apparent_crossvein_only,absent,,,,,
Forewing  second branch of radial sector  RS2  vein presence,present,absent,,,,,,
Forewing  anal crossvein  a  presence,present,absent,,,,,,
Forewing  nygmata  sensory spots  on wing membrane  in second submarginal cell ,present,absent,,,,,,
Hind wing  costal  C  vein presence,"present, and at least half as long as distance to costal break (stigma)",absent_or_reduced_to_a_small_stub_at_wing_base,,,,,,
Hind wing  non apical hamuli  presence,present_and_hooked_or_thick_and_highly_sclerotized,absent,,,,,,
Hind wing  subcostal  SC  vein presence,present,absent,,,,,,
HInd wing  number of rs m crossveins,3,1,_or_0,,,,,
Hind wing  m cu crossvein presence,present,absent,,,,,,
Hind wing  medial  M  vein and cubital  Cu  vein fusion,separated_basally,1,_or_0,,,,,
Hind wing  costal margin,not_broadly_curved,broadly_curved,,,,,,
Hind wing  costal trough extending from base to hamuli,present,absent,,,,,,
Hind wing  fusion of radial  R  and costal  C  veins posterad hamuli  cell presence,"R and C separated and forming a small cell posterad hamuli separating and re-fusing distal to hamuli, thus forming a small cel","R and C not separated posterior to hamuli, no small cell  present",,,,,,
Hind wing  microspicules posterad hamuli,present,absent,,,,,,
Hind wing  presence of guide setae or spines posterior to hamuli,absent,present,,,,,,
Hind wing  presence of sclerotized glabrous plate posterior to hamuli,absent,present,,,,,,
Mesotibial apical spurs,two,one,none,,,,,
Tip of tibial spurs,pointed and sclerotized,blunt and with a smooth (membranous) area,,,,,,
Presence of velum on calcar,velum present,velum absent,,,,,,
Velum appearance,smooth and with smooth rim,smooth but with serrated rim,smooth but with a fine comb of socketed setae along edge,with a spinose or ribbed surface,,,,
Anterior basitarsus,with one row of paddle-shaped setae,with more than one row of paddle-shaped setae,with ordinary setae,without paddle-shaped setae though sometimes with blunt or filiform setae,without setae,,,
Inner posterior metatibial spur,simple,modified into a calcar,,,,,,
Mesotibial apical sensillum,present,absent,,,,,,
Inner posterior mesotibial spur,simple,modified into a calcar,,,,,,
Foretibial apical sensillum,present,absent,,,,,,
Metatibial apical sensillum,present,absent,,,,,,
Midleg tibial comb,absent,"present as row of similarly sized, peg-like setae arranged in a row",,,,,,
Abdominal tergum 3  metasomal 2  presence of laterotergites,not separated by a longitudinal fold into lateral and medial tergites,separated by a longitudinal fold into lateral and medial tergites,,,,,,
Interleaving of abdominal terga and sterna 4 5  metasomal terga 3 4 ,"not interleaved. If in contact, terga completely overlying preceding sterna",abdominal tergum 4 lying under sternum 3 laterally (metasomal tergum 3 under sternum 2),,,,,,
Presence of glymma  laterope ,absent,present,,,,,,
Articulation between abdominal tergum 2 and tergum 3  metasomal 1 and 2 ,terga 2-3 not hinged sub-medially,terga 2-3 hinged submedially,,,,,,
Spiracles on abdominal terga 2 8  metasomal 1 7 ,well_developed_on_most_terga,"reduced or absent on 2-7, present or absent on 8",,,,,,
Second abdominal sternum  first metasomal ,entirely_evenly_sclerotized,"evenly sclerotized anteriorly, but posteriorly with membranous areas, or more weakly sclerotized areas",,,,,,
Abdominal sternum 1  presence,present,absent,,,,,,
Abdominal tergum 2  metasomal 1  overlapping abdominal tergum 3  metasomal 2 ,overlapping,not_overlapping,,,,,,
Male thoracic muscles diploid,haploid,diploid,,,,,,
Position of egg exit,at or near apex of ovipositor,at or near base of ovipositor,,,,,,
Ovipositor shape,laterally flattened,"cylindrical, not laterally flattened",,,,,,
Furcula of ovipositor,absent,present,,,,,,
Structure of ovipositor sheaths,unisegmented,divided into two segments,,,,,,
Valvilli presence,absent,present,,,,,,
Second valvifer structure,"not divided, pars articularis absent","subdivided, pars articularis present",,,,,,
Meconium,"not formed, feces voided continuously during larval development",meconium voided at the end of larval development,,,,,,
Mode of parasitism,ectoparasitoids,endoparasitoids,,,,,,
Larval food source,plants,animals,,,,,,
Silk production,labial silk production from salivary gland secretions,no silk production from salivary gland secretions,,,,,,
